A few points...
High voltages rarely ever harm tubes, despite what the specs say. Excessive power dissipation is what damages tubes or shortens their lives, whether anode or screen grid dissipation. The main danger with high voltages is the potential to cause excessive power dissipation. There is ultimately a point where arc-over starts to happen inside a tube, but you have to go to real stupid high voltages for that.
It is essential to have screen grid protection resistors, when you have a screen grid supply with a filter capacitor (like in most amps). This is because when the amp cranks, the filter cap will dump a load of current into the screen grids, so they go to the edge of melt-down, especially on transients. A lot of vintage amps use a 100 ohm value, but IMO that's too low if you want to preserve your tubes. I would never use less than 1k with any EL84 amp. See the comments in this EL84 review by Randall Aiken -
http://www.aikenamps.com/JJ_EH_tube_tests.html I've also just finished repairing an amp for a customer, which had 100 ohm screen resistors. The EL84s were pretty much ruined and one was red-plating. Needless to say I put in 1k screen protection resistors to help protect the new set.
A choke also has a current limiting effect and can help protect screen grids. The fundamental action of a choke is to fight changes in current through the choke. So when screen grids start to draw more current on a transient, the choke will rapidly drop the voltage seen at the screen grids. The result is to make an amp compress more, with a softer attack and a looser feel - like a Vox AC30. Larger screen resistor values also make for more compression, although not as much typically as with a choke. Just for refernce you may like to compare these schematics from an AC30 and a Matchless Lightning with the TC-15's:
http://www.schematicheaven.com/voxamps/ ... weramp.pdfhttp://www.schematicheaven.com/newamps/ ... htning.pdfThere is no reason for an amp to sound brittle or harsh with either a choke or adequately sized screen resistors. There are thousands of EL84 amps out there which have awesome tone, and which aren't melting down their screen grids. One of the amps I often gig with has 1k3 resistors going to each screen grid (just what I had handy at the time), and has a tone as smooth as butter. If an amp sounds harsh or brittle once you've put in sensible screen grid protection measures, then that indicates that a problem elsewhere in the amp has been unmasked. Possibly even just poor quality power tubes.
There are inevitably a lot of subjective opinions about what screen protection resistor value sounds best. I've even come across one guy who swears by his 2k7 screen grid resistors. He's an experienced blues player, and I so don't argue with him.
