Very good trouble shooting pointers from Coco. Particularly the click/pop tests.
Yes, for the plates set your meter to DC and use the red probe, black to ground.
It doesn't matter which speaker output you plug into, unless it's wired wrong
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2.6-6.0v on the plates is definitely far too low. Just to be sure, that's the range you are getting on pins 1 and 6 of the preamp tubes and pin 3 of the power tubes? I would be checking the wiring for something shorted.The fuse should have blown but it sounds like a short somewhere. It could just be something mis-wired though. You said you have 700 at the PT secondaries so that's good. Check backwards from there to see where you lose the voltage. What are you getting on pin 3 of the output tubes? What about pin 1 and 6 of the phase inverter and the preamp tubes? Again, you can check this with tubes out, the voltages won't match the schematic (without the tubes the voltages should be higher) but you just want to find where it disappears or goes very low. Wherever you lose the voltage, or it goes low, that's where your problem is.
It's not uncommon to have more than one issue on a first build. If you find one issue and fixing it doesn't resolve the issues, or seems to cause another issue, continue trouble shooting from where you fixed the first problem.
Remember, there *should* be very high voltages on the plates, keep your left hand behind your back and use a pointed probe, not clips, to check the voltages. Don't touch anything with your bare hands while poking around in a live amp. Wear rubber soled shoes as well, especially if you are on a concrete floor.