The two 470k resistors (R8 and R42) on the MkII form what is called a "voltage divider." It is a circuit that reduces the amount of voltage passing through it. In this case, where the two resistors are of equal value, it divides the voltage in half. The result in this application is that there would be less gain headed into V2A because this voltage divider cuts it in half.
If you want a
really good series of videos on everything resistors do in a tube amp (including voltage dividers) then I
highly recommend Uncle Doug's five-part video series starting with part one here....
https://www.youtube.com/watch?v=LBtoarc6NlsIt starts pretty basic but gets very in-depth and tube-circuit specific before too long.